Comma

To quote from elsewhere:-
"This butterfly was once widespread over most of England and Wales, and parts of southern Scotland, but by the middle of the 1800s had suffered a severe decline that left it confined to the Welsh border counties, especially West Gloucestershire, East Gloucestershire, Herefordshire and Monmouthshire. It is thought that the decline may have been due to a reduction in Hop farming, a key larval foodplant at the time. Since the 1960s this butterfly has made a spectacular comeback, with a preference for Common Nettle as the larval foodplant, and it is now found throughout England, Wales, the Isle of Man and the Channel Islands and has recently reached Scotland."